(1) DP1*@cell+p14ARF*@nucleus -> DP1*:p14ARF*@nucleus (2) E2F1@nucleus+p14ARF*@nucleus -> E2F1:p14ARF*@nucleus (3) E2F1@nucleus - p14ARF*@nucleus -> (4) MDM2@nucleus+p14ARF*@nucleus -> MDM2:p14ARF*@nucleus